How to use cucumber-jvm without lein-cucumber

features.clj

(ns cucutest.features
  (:require [clojure.test :refer :all])
  (:import [cucumber.api.cli Main]))

(deftest ^:cucumber run-cucumber
  (let [cucumber-args ["--plugin" "progress" "--glue" "test/features/step_definitions" "test/features"]]
    (Main/main (into-array cucumber-args))))

project.clj

;; given the features in test/features
;; and the step definitions in test/features/step_definitions
;; lein pep will execute the features in the test/features directory
;; via the test in test/cucutest/features.clj file
;; which is marked as :cucumber

(defproject cucutest "0.1.0-SNAPSHOT"
  :description "FIXME: write description"
  :url "http://example.com/FIXME"
  :license {:name "Eclipse Public License"
            :url "http://www.eclipse.org/legal/epl-v10.html"}
  :dependencies [[org.clojure/clojure "1.8.0"]]
  :profiles {:dev {:dependencies [[info.cukes/cucumber-clojure "1.2.4"]]}
             :cucumber-run {:test-paths ["test/features/step_definitions"]}}
  :test-selectors {:default (complement :cucumber)
                   :cucumber :cucumber
                   :all (constantly true)}
  :aliases {"pep" ["with-profile" "+cucumber-run" "test" ":cucumber"]})