Protocol functions fail to find extensions is assigned to locals or other var

protocol-function-failures.clj

(defprotocol TestProtocol
  (tester [o]))

(let [t tester]
  (defn another-tester [o]
  	(t o)))

(def another-tester2 tester)

(extend-protocol TestProtocol
  String
  (tester [o] (println "Strings work!")))

(another-tester "A") ;; Error
(another-tester2 "A") ;; Error
(tester "A") ;; Works fine

(let [t tester]
  (defn another-tester [o]
  	(t o)))

(another-tester "A") ;; Works fine

(def another-tester2 tester)

(another-tester2 "A") ;; Works fine

(extend-protocol TestProtocol
  Long
  (tester [o] (println "Longs work!")))

(another-tester "A") ;; Works fine
(another-tester 3) ;; Error
(another-tester2 3) ;; Error